Professor Colin Torney
- Professor of Applied Mathematics (Mathematics)
                        email: 
                        Colin.Torney@glasgow.ac.uk
                    
Mathematics Building
Research interests
My research uses mathematical modelling and machine learning to study ecology and animal movement.
Please see my website for further information:
https://www.maths.gla.ac.uk/~ctorney/
Research groups
Publications
2025
    
    Kundegorski, Mikolaj E., Honkanen, Hannele M., Stephen, Alastair, Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Killen, Shaun 
ORCID: https://orcid.org/0000-0003-4949-3988 and Adams, Colin E. 
ORCID: https://orcid.org/0000-0003-2470-9754
  
(2025)
  
Defining the water flow cues for navigation in migrating Atlantic salmon smolts.
    
    Journal of Fish Biology, 107(1),
	  pp. 161-168.
     
    
    (doi: 10.1111/jfb.70004)
    (PMID:40070053) 
     
  
  
    
    Campioni, Nazareno, Husmeier, Dirk ORCID: https://orcid.org/0000-0003-1673-7413, Gaskell, Jennifer 
ORCID: https://orcid.org/0000-0001-9583-323X, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490 and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2025)
  
Scalable non-Markovian state switching models for animal movement.
    
    
      In: Giuggioli, Luca and Maini, Philip (eds.)
      The Mathematics of Movement: an Interdisciplinary Approach to Mutual Challenges in Animal Ecology and Cell Biology.
    
    
    Springer Nature.
        
        
  
   (Accepted for Publication)
    
    Kavwele, Cyrus M., Hopcraft, J. Grant C. ORCID: https://orcid.org/0000-0002-9617-9951, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490, Nyafi, Gerald, Kimuya, Nancy and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2025)
  
Real‐time classification of Serengeti wildebeest behaviour with edge machine learning and a long‐range IoT network.
    
    Canadian Journal of Zoology, 103,
	  pp. 1-11.
     
    
    (doi: 10.1139/cjz-2024-0075)
     
     
  
  
2024
    
    Masolele, Majaliwa M., Hopcraft, J. Grant C. ORCID: https://orcid.org/0000-0002-9617-9951 and Torney, Colin J.T. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2024)
  
Efficient approximate Bayesian inference for quantifying uncertainty in multiscale animal movement models.
    
    Ecological Informatics, 84,
	 
      102853.
    
    (doi: 10.1016/j.ecoinf.2024.102853)
     
     
  
  
    
    Kavwele, Cyrus M., Hopcraft, J. Grant C. ORCID: https://orcid.org/0000-0002-9617-9951, Davy, Deborah and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2024)
  
Automated and repeated aerial observations of GPS‐collared animals using UAVs and open‐source electronics.
    
    Ecosphere, 15(4),
	 
      e4841.
    
    (doi: 10.1002/ecs2.4841)
     
     
  
  
2023
    
    Rathore, Akanksha, Sharma, Ananth, Shah, Shaan, Sharma, Nitika, Torney, Colin ORCID: https://orcid.org/0000-0003-1673-7835 and Guttal, Vishwesha
  
(2023)
  
Multi-Object Tracking in Heterogeneous environments (MOTHe) for animal video recordings.
    
    PeerJ, 11,
	 
      e15573.
    
    (doi: 10.7717/peerj.15573)
    (PMID:37397020) 
    (PMCID:PMC10309051) 
  
  
    
    Paun, Ionut ORCID: https://orcid.org/0009-0003-1673-9803, Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413 and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2023)
  
Stochastic variational inference for scalable non-stationary Gaussian process regression.
    
    Statistics and Computing, 33(2),
	 
      44.
    
    (doi: 10.1007/s11222-023-10210-w)
     
     
  
  
    
    Gaskell, Jennifer ORCID: https://orcid.org/0000-0001-9583-323X, Campioni, Nazareno, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490, Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413 and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2023)
  
Inferring the interaction rules of complex systems with graph neural networks and approximate Bayesian computation.
    
    Journal of the Royal Society: Interface, 20(198),
	 
      20220676.
    
    (doi: 10.1098/rsif.2022.0676)
    (PMID:36596456) 
     
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Laxton, Megan, Lloyd‐Jones, David J., Kohi, Edward M., Frederick, Howard L., Moyer, David C., Mrisha, Chediel, Mwita, Machoke and Hopcraft, J. Grant C. 
ORCID: https://orcid.org/0000-0002-9617-9951
  
(2023)
  
Estimating the abundance of a group-living species using multi-latent spatial models.
    
    Methods in Ecology and Evolution, 14(1),
	  pp. 77-86.
     
    
    (doi: 10.1111/2041-210x.13941)
     
     
  
  
2022
    
    Paun, Ionut ORCID: https://orcid.org/0009-0003-1673-9803, Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413, Hopcraft, J. Grant C. 
ORCID: https://orcid.org/0000-0002-9617-9951, Masolele, Majaliwa M. and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2022)
  
Inferring spatially varying animal movement characteristics using a hierarchical continuous-time velocity model.
    
    Ecology Letters, 25(12),
	  pp. 2726-2738.
     
    
    (doi: 10.1111/ele.14117)
    (PMID:36256526) 
     
  
  
    
    Kavwele, Cyrus M., Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Morrison, Thomas A. 
ORCID: https://orcid.org/0000-0001-7262-1402, Fulford, Sidney, Masolele, Majaliwa M., Masoy, Joseph and Hopcraft, J. Grant C. 
ORCID: https://orcid.org/0000-0002-9617-9951
  
(2022)
  
Non‐local effects of human activity on the spatial distribution of migratory wildlife in Serengeti National Park, Tanzania.
    
    Ecological Solutions and Evidence, 3(3),
	 
      e12159.
    
    (doi: 10.1002/2688-8319.12159)
     
     
  
  
2021
    
    Campioni, Nazareno, Husmeier, Dirk ORCID: https://orcid.org/0000-0003-1673-7413, Morales, Juan 
ORCID: https://orcid.org/0000-0001-7269-7490, Gaskell, Jennifer 
ORCID: https://orcid.org/0000-0001-9583-323X and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2021)
  
Inferring microscale properties of interacting systems from macroscale observations.
    
    Physical Review Research, 3(4),
	 
      043074.
    
    (doi: 10.1103/PhysRevResearch.3.043074)
     
     
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490 and Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413
  
(2021)
  
A hierarchical machine learning framework for the analysis of large scale animal movement data.
    
    Movement Ecology, 9,
	 
      6.
    
    (doi: 10.1186/s40462-021-00242-0)
    (PMID:33602302) 
     
  
  
2020
    
    Campioni, Nazareno, Husmeier, Dirk ORCID: https://orcid.org/0000-0003-1673-7413, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490, Gaskell, Jennifer 
ORCID: https://orcid.org/0000-0001-9583-323X and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2020)
  
Modelling multiscale collective behavior with Gaussian processes.
    
    
      In: Ladde, Gangaram and Samia, Noelle (eds.)
      Proceedings of the 2nd International Conference on Statistics: Theory and Applications (ICSTA'20).
    
    
    Avestia Publishing: Ottawa, Canada, p. 124.
         ISBN 9781927877685
        (doi: 10.11159/icsta20.124)
  
  
    
    Gaskell, Jennifer ORCID: https://orcid.org/0000-0001-9583-323X, Campioni, Nazareno, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490, Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413 and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2020)
  
Approximate Bayesian inference for individual-based models with emergent dynamics.
    
    
      In: Ladde, Gangaram and Samia, Noelle (eds.)
      Proceedings of the 2nd International Conference on Statistics: Theory and Applications (ICSTA'20).
    
    
    Avestia Publishing: Ottawa, Canada, p. 125.
         ISBN 9781927877685
        (doi: 10.11159/icsta20.125)
  
  
2019
    
    Evans, Julian C., Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Votier, Stephen C. and Dall, Sasha R. X.
  
(2019)
  
Social information use and collective foraging in a pursuit diving seabird.
    
    PLoS ONE, 14(9),
	 
      e0222600.
    
    (doi: 10.1371/journal.pone.0222600)
    (PMID:31545848) 
    (PMCID:PMC6756525) 
  
  
    
    Paun, Ionut ORCID: https://orcid.org/0009-0003-1673-9803, Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413 and Torney, Colin 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2019)
  
A Study on Discrete-Time Movement Models.
    
    In: International Conference on Statistics: Theory and Applications (ICSTA’19), Lisbon, Portugal, 13-14 Aug 2019,
     p. 27. 
     ISBN 9781927877647
    (doi: 10.11159/icsta19.27)
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Lloyd-Jones, David J., Chevallier, Mark, Moyer, David C., Maliti, Honori T., Mwita, Machoke, Kohi, Edward M. and Hopcraft, J.G.C. 
ORCID: https://orcid.org/0000-0002-9617-9951
  
(2019)
  
A comparison of deep learning and citizen science techniques for counting wildlife in aerial survey images.
    
    Methods in Ecology and Evolution, 10(6),
	  pp. 779-787.
     
    
    (doi: 10.1111/2041-210X.13165)
     
     
  
  
2018
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Hopcraft, J. Grant C. 
ORCID: https://orcid.org/0000-0002-9617-9951, Morrison, Thomas A. 
ORCID: https://orcid.org/0000-0001-7262-1402, Couzin, Iain D. and Levin, Simon A.
  
(2018)
  
From single steps to mass migration: the problem of scale in the movement ecology of the Serengeti wildebeest.
    
    Philosophical Transactions of the Royal Society B: Biological Sciences, 373(1746),
	 
      20170012.
    
    (doi: 10.1098/rstb.2017.0012)
    (PMID:29581397) 
    (PMCID:PMC5882982) 
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Lamont, Myles, Debell, Leon, Angohiatok, Ryan J., Leclerc, Lisa-Marie and Berdahl, Andrew M.
  
(2018)
  
Inferring the rules of social interaction in migrating caribou.
    
    Philosophical Transactions of the Royal Society B: Biological Sciences, 373(1746),
	 
      20170385.
    
    (doi: 10.1098/rstb.2017.0385)
    (PMID:29581404) 
     
  
  
    
    Westley, Peter A.H., Berdahl, Andrew M., Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835 and Biro, Dora
  
(2018)
  
Collective movement in ecology: from emerging technologies to conservation and management.
    
    Philosophical Transactions of the Royal Society B: Biological Sciences, 373(1746),
	 
      20170004.
    
    (doi: 10.1098/rstb.2017.0004)
    (PMID:29581389) 
     
  
  
    
    Faust, Christina L. ORCID: https://orcid.org/0000-0002-8824-7424, McCallum, Hamish I., Bloomfield, Laura S.P., Gottdenker, Nicole L., Gillespie, Thomas R., Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835, Dobson, Andrew P. and Plowright, Raina K.
  
(2018)
  
Pathogen spillover during land conversion.
    
    Ecology Letters, 21(4),
	  pp. 471-483.
     
    
    (doi: 10.1111/ele.12904)
    (PMID:29466832) 
     
  
  
2017
    
    Ioannou, Christos C., Ramnarine, Indar W. and Torney, Colin ORCID: https://orcid.org/0000-0003-1673-7835
  
(2017)
  
High-predation habitats affect the social dynamics of collective exploration in a shoaling fish.
    
    Science Advances, 3(5),
	 
      e1602682.
    
    (doi: 10.1126/sciadv.1602682)
     
     
  
  
    
    Valletta, John Joseph, Torney, Colin ORCID: https://orcid.org/0000-0003-1673-7835, Kings, Michael, Thornton, Alex and Madden, Joah
  
(2017)
  
Applications of machine learning in animal behaviour studies.
    
    Animal Behaviour, 124,
	  pp. 203-220.
     
    
    (doi: 10.1016/j.anbehav.2016.12.005)
     
     
  
  
2016
    
    Berdahl, Andrew, van Leeuwen, Anieke, Levin, Simon A. and Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835
  
(2016)
  
Collective behavior as a driver of critical transitions in migratory populations.
    
    Movement Ecology, 4(1),
	 
      18.
    
    (doi: 10.1186/s40462-016-0083-8)
    (PMID:27429757) 
    (PMCID:PMC4946155) 
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Dobson, Andrew P., Borner, Felix, Lloyd-Jones, David J., Moyer, David, Maliti, Honori T., Mwita, Machoke, Fredrick, Howard, Borner, Markus and Hopcraft, J. Grant C. 
ORCID: https://orcid.org/0000-0002-9617-9951
  
(2016)
  
Assessing rotation-invariant feature classification for automated wildebeest population counts.
    
    PLoS ONE, 11(5),
	 
      e0156342.
    
    (doi: 10.1371/journal.pone.0156342)
    (PMID:27227888) 
    (PMCID:PMC4881999) 
  
  
2015
    
    Hein, Andrew M., Rosenthal, Sara Brin, Hagstrom, George I., Berdahl, Andrew, Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835 and Couzin, Iain D.
  
(2015)
  
The evolution of distributed sensing and collective computation in animal populations.
    
    eLife, 4,
	 
      e10955.
    
    (doi: 10.7554/eLife.10955)
    (PMID:26652003) 
    (PMCID:PMC4755780) 
  
  
    
    Berdahl, Andrew, Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Schertzer, Emmanuel and Levin, Simon A.
  
(2015)
  
On the evolutionary interplay between dispersal and local adaptation in heterogeneous environments.
    
    Evolution, 69(6),
	  pp. 1390-1405.
     
    
    (doi: 10.1111/evo.12664)
     
     
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Lorenzi, Tommaso, Couzin, Iain D. and Levin, Simon A.
  
(2015)
  
Social information use and the evolution of unresponsiveness in collective systems.
    
    Journal of the Royal Society: Interface, 12(103),
	 
     
    
    (doi: 10.1098/rsif.2014.0893)
    (PMID:25519991) 
    (PMCID:PMC4305405) 
  
  
2014
    
    Kao, Albert B., Miller, Noam, Torney, Colin ORCID: https://orcid.org/0000-0003-1673-7835, Hartnett, Andrew and Couzin, Iain D.
  
(2014)
  
Collective learning and optimal consensus decisions in social animal groups.
    
    PLoS Computational Biology, 10(8),
	 
      e1003762.
    
    (doi: 10.1371/journal.pcbi.1003762)
    (PMID:25101642) 
    (PMCID:PMC4125046) 
  
  
2013
Strandburg-Peshkin, A. et al. (2013) Visual sensory networks and effective information transfer in animal groups. Current Biology, 23(17), R709-R711. (doi: 10.1016/j.cub.2013.07.059) (PMID:24028946) (PMCID:PMC4780851)
    
    Coburn, Luke, Cerone, Luca, Torney, Colin ORCID: https://orcid.org/0000-0003-1673-7835, Couzin, Iain D and Neufeld, Zoltan
  
(2013)
  
Tactile interactions lead to coherent motion and enhanced chemotaxis of migrating cells.
    
    Physical Biology, 10(4),
	 
      046002.
    
    (doi: 10.1088/1478-3975/10/4/046002)
    (PMID:23752100) 
     
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Levin, Simon A. and Couzin, Iain D.
  
(2013)
  
Decision accuracy and the role of spatial interaction in opinion dynamics.
    
    Journal of Statistical Physics, 151(1-2),
	  pp. 203-217.
     
    
    (doi: 10.1007/s10955-013-0700-5)
     
     
  
  
    
    Berdahl, Andrew, Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Ioannou, Christos C., Faria, Jolyon J. and Couzin, Iain D.
  
(2013)
  
Emergent sensing of complex environments by mobile animal groups.
    
    Science, 339(6119),
	  pp. 574-576.
     
    
    (doi: 10.1126/science.1225883)
    (PMID:23372013) 
     
  
  
2011
    
    Couzin, Iain D., Ioannou, Christos C., Demirel, Güven, Gross, Thilo, Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Hartnett, Andrew, Conradt, Larissa, Levin, Simon A. and Leonard, Naomi E.
  
(2011)
  
Uninformed individuals promote democratic consensus in animal groups.
    
    Science, 334(6062),
	  pp. 1578-1580.
     
    
    (doi: 10.1126/science.1210280)
    (PMID:22174256) 
     
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Berdahl, Andrew and Couzin, Iain D.
  
(2011)
  
Signalling and the evolution of cooperative foraging in dynamic environments.
    
    PLoS Computational Biology, 7(9),
	  e1002194.
     
    
    (doi: 10.1371/journal.pcbi.1002194)
    (PMID:21966265) 
    (PMCID:PMC3178622) 
  
  
Articles
    
    Kundegorski, Mikolaj E., Honkanen, Hannele M., Stephen, Alastair, Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Killen, Shaun 
ORCID: https://orcid.org/0000-0003-4949-3988 and Adams, Colin E. 
ORCID: https://orcid.org/0000-0003-2470-9754
  
(2025)
  
Defining the water flow cues for navigation in migrating Atlantic salmon smolts.
    
    Journal of Fish Biology, 107(1),
	  pp. 161-168.
     
    
    (doi: 10.1111/jfb.70004)
    (PMID:40070053) 
     
  
  
    
    Kavwele, Cyrus M., Hopcraft, J. Grant C. ORCID: https://orcid.org/0000-0002-9617-9951, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490, Nyafi, Gerald, Kimuya, Nancy and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2025)
  
Real‐time classification of Serengeti wildebeest behaviour with edge machine learning and a long‐range IoT network.
    
    Canadian Journal of Zoology, 103,
	  pp. 1-11.
     
    
    (doi: 10.1139/cjz-2024-0075)
     
     
  
  
    
    Masolele, Majaliwa M., Hopcraft, J. Grant C. ORCID: https://orcid.org/0000-0002-9617-9951 and Torney, Colin J.T. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2024)
  
Efficient approximate Bayesian inference for quantifying uncertainty in multiscale animal movement models.
    
    Ecological Informatics, 84,
	 
      102853.
    
    (doi: 10.1016/j.ecoinf.2024.102853)
     
     
  
  
    
    Kavwele, Cyrus M., Hopcraft, J. Grant C. ORCID: https://orcid.org/0000-0002-9617-9951, Davy, Deborah and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2024)
  
Automated and repeated aerial observations of GPS‐collared animals using UAVs and open‐source electronics.
    
    Ecosphere, 15(4),
	 
      e4841.
    
    (doi: 10.1002/ecs2.4841)
     
     
  
  
    
    Rathore, Akanksha, Sharma, Ananth, Shah, Shaan, Sharma, Nitika, Torney, Colin ORCID: https://orcid.org/0000-0003-1673-7835 and Guttal, Vishwesha
  
(2023)
  
Multi-Object Tracking in Heterogeneous environments (MOTHe) for animal video recordings.
    
    PeerJ, 11,
	 
      e15573.
    
    (doi: 10.7717/peerj.15573)
    (PMID:37397020) 
    (PMCID:PMC10309051) 
  
  
    
    Paun, Ionut ORCID: https://orcid.org/0009-0003-1673-9803, Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413 and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2023)
  
Stochastic variational inference for scalable non-stationary Gaussian process regression.
    
    Statistics and Computing, 33(2),
	 
      44.
    
    (doi: 10.1007/s11222-023-10210-w)
     
     
  
  
    
    Gaskell, Jennifer ORCID: https://orcid.org/0000-0001-9583-323X, Campioni, Nazareno, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490, Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413 and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2023)
  
Inferring the interaction rules of complex systems with graph neural networks and approximate Bayesian computation.
    
    Journal of the Royal Society: Interface, 20(198),
	 
      20220676.
    
    (doi: 10.1098/rsif.2022.0676)
    (PMID:36596456) 
     
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Laxton, Megan, Lloyd‐Jones, David J., Kohi, Edward M., Frederick, Howard L., Moyer, David C., Mrisha, Chediel, Mwita, Machoke and Hopcraft, J. Grant C. 
ORCID: https://orcid.org/0000-0002-9617-9951
  
(2023)
  
Estimating the abundance of a group-living species using multi-latent spatial models.
    
    Methods in Ecology and Evolution, 14(1),
	  pp. 77-86.
     
    
    (doi: 10.1111/2041-210x.13941)
     
     
  
  
    
    Paun, Ionut ORCID: https://orcid.org/0009-0003-1673-9803, Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413, Hopcraft, J. Grant C. 
ORCID: https://orcid.org/0000-0002-9617-9951, Masolele, Majaliwa M. and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2022)
  
Inferring spatially varying animal movement characteristics using a hierarchical continuous-time velocity model.
    
    Ecology Letters, 25(12),
	  pp. 2726-2738.
     
    
    (doi: 10.1111/ele.14117)
    (PMID:36256526) 
     
  
  
    
    Kavwele, Cyrus M., Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Morrison, Thomas A. 
ORCID: https://orcid.org/0000-0001-7262-1402, Fulford, Sidney, Masolele, Majaliwa M., Masoy, Joseph and Hopcraft, J. Grant C. 
ORCID: https://orcid.org/0000-0002-9617-9951
  
(2022)
  
Non‐local effects of human activity on the spatial distribution of migratory wildlife in Serengeti National Park, Tanzania.
    
    Ecological Solutions and Evidence, 3(3),
	 
      e12159.
    
    (doi: 10.1002/2688-8319.12159)
     
     
  
  
    
    Campioni, Nazareno, Husmeier, Dirk ORCID: https://orcid.org/0000-0003-1673-7413, Morales, Juan 
ORCID: https://orcid.org/0000-0001-7269-7490, Gaskell, Jennifer 
ORCID: https://orcid.org/0000-0001-9583-323X and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2021)
  
Inferring microscale properties of interacting systems from macroscale observations.
    
    Physical Review Research, 3(4),
	 
      043074.
    
    (doi: 10.1103/PhysRevResearch.3.043074)
     
     
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490 and Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413
  
(2021)
  
A hierarchical machine learning framework for the analysis of large scale animal movement data.
    
    Movement Ecology, 9,
	 
      6.
    
    (doi: 10.1186/s40462-021-00242-0)
    (PMID:33602302) 
     
  
  
    
    Evans, Julian C., Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Votier, Stephen C. and Dall, Sasha R. X.
  
(2019)
  
Social information use and collective foraging in a pursuit diving seabird.
    
    PLoS ONE, 14(9),
	 
      e0222600.
    
    (doi: 10.1371/journal.pone.0222600)
    (PMID:31545848) 
    (PMCID:PMC6756525) 
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Lloyd-Jones, David J., Chevallier, Mark, Moyer, David C., Maliti, Honori T., Mwita, Machoke, Kohi, Edward M. and Hopcraft, J.G.C. 
ORCID: https://orcid.org/0000-0002-9617-9951
  
(2019)
  
A comparison of deep learning and citizen science techniques for counting wildlife in aerial survey images.
    
    Methods in Ecology and Evolution, 10(6),
	  pp. 779-787.
     
    
    (doi: 10.1111/2041-210X.13165)
     
     
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Hopcraft, J. Grant C. 
ORCID: https://orcid.org/0000-0002-9617-9951, Morrison, Thomas A. 
ORCID: https://orcid.org/0000-0001-7262-1402, Couzin, Iain D. and Levin, Simon A.
  
(2018)
  
From single steps to mass migration: the problem of scale in the movement ecology of the Serengeti wildebeest.
    
    Philosophical Transactions of the Royal Society B: Biological Sciences, 373(1746),
	 
      20170012.
    
    (doi: 10.1098/rstb.2017.0012)
    (PMID:29581397) 
    (PMCID:PMC5882982) 
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Lamont, Myles, Debell, Leon, Angohiatok, Ryan J., Leclerc, Lisa-Marie and Berdahl, Andrew M.
  
(2018)
  
Inferring the rules of social interaction in migrating caribou.
    
    Philosophical Transactions of the Royal Society B: Biological Sciences, 373(1746),
	 
      20170385.
    
    (doi: 10.1098/rstb.2017.0385)
    (PMID:29581404) 
     
  
  
    
    Westley, Peter A.H., Berdahl, Andrew M., Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835 and Biro, Dora
  
(2018)
  
Collective movement in ecology: from emerging technologies to conservation and management.
    
    Philosophical Transactions of the Royal Society B: Biological Sciences, 373(1746),
	 
      20170004.
    
    (doi: 10.1098/rstb.2017.0004)
    (PMID:29581389) 
     
  
  
    
    Faust, Christina L. ORCID: https://orcid.org/0000-0002-8824-7424, McCallum, Hamish I., Bloomfield, Laura S.P., Gottdenker, Nicole L., Gillespie, Thomas R., Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835, Dobson, Andrew P. and Plowright, Raina K.
  
(2018)
  
Pathogen spillover during land conversion.
    
    Ecology Letters, 21(4),
	  pp. 471-483.
     
    
    (doi: 10.1111/ele.12904)
    (PMID:29466832) 
     
  
  
    
    Ioannou, Christos C., Ramnarine, Indar W. and Torney, Colin ORCID: https://orcid.org/0000-0003-1673-7835
  
(2017)
  
High-predation habitats affect the social dynamics of collective exploration in a shoaling fish.
    
    Science Advances, 3(5),
	 
      e1602682.
    
    (doi: 10.1126/sciadv.1602682)
     
     
  
  
    
    Valletta, John Joseph, Torney, Colin ORCID: https://orcid.org/0000-0003-1673-7835, Kings, Michael, Thornton, Alex and Madden, Joah
  
(2017)
  
Applications of machine learning in animal behaviour studies.
    
    Animal Behaviour, 124,
	  pp. 203-220.
     
    
    (doi: 10.1016/j.anbehav.2016.12.005)
     
     
  
  
    
    Berdahl, Andrew, van Leeuwen, Anieke, Levin, Simon A. and Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835
  
(2016)
  
Collective behavior as a driver of critical transitions in migratory populations.
    
    Movement Ecology, 4(1),
	 
      18.
    
    (doi: 10.1186/s40462-016-0083-8)
    (PMID:27429757) 
    (PMCID:PMC4946155) 
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Dobson, Andrew P., Borner, Felix, Lloyd-Jones, David J., Moyer, David, Maliti, Honori T., Mwita, Machoke, Fredrick, Howard, Borner, Markus and Hopcraft, J. Grant C. 
ORCID: https://orcid.org/0000-0002-9617-9951
  
(2016)
  
Assessing rotation-invariant feature classification for automated wildebeest population counts.
    
    PLoS ONE, 11(5),
	 
      e0156342.
    
    (doi: 10.1371/journal.pone.0156342)
    (PMID:27227888) 
    (PMCID:PMC4881999) 
  
  
    
    Hein, Andrew M., Rosenthal, Sara Brin, Hagstrom, George I., Berdahl, Andrew, Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835 and Couzin, Iain D.
  
(2015)
  
The evolution of distributed sensing and collective computation in animal populations.
    
    eLife, 4,
	 
      e10955.
    
    (doi: 10.7554/eLife.10955)
    (PMID:26652003) 
    (PMCID:PMC4755780) 
  
  
    
    Berdahl, Andrew, Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Schertzer, Emmanuel and Levin, Simon A.
  
(2015)
  
On the evolutionary interplay between dispersal and local adaptation in heterogeneous environments.
    
    Evolution, 69(6),
	  pp. 1390-1405.
     
    
    (doi: 10.1111/evo.12664)
     
     
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Lorenzi, Tommaso, Couzin, Iain D. and Levin, Simon A.
  
(2015)
  
Social information use and the evolution of unresponsiveness in collective systems.
    
    Journal of the Royal Society: Interface, 12(103),
	 
     
    
    (doi: 10.1098/rsif.2014.0893)
    (PMID:25519991) 
    (PMCID:PMC4305405) 
  
  
    
    Kao, Albert B., Miller, Noam, Torney, Colin ORCID: https://orcid.org/0000-0003-1673-7835, Hartnett, Andrew and Couzin, Iain D.
  
(2014)
  
Collective learning and optimal consensus decisions in social animal groups.
    
    PLoS Computational Biology, 10(8),
	 
      e1003762.
    
    (doi: 10.1371/journal.pcbi.1003762)
    (PMID:25101642) 
    (PMCID:PMC4125046) 
  
  
Strandburg-Peshkin, A. et al. (2013) Visual sensory networks and effective information transfer in animal groups. Current Biology, 23(17), R709-R711. (doi: 10.1016/j.cub.2013.07.059) (PMID:24028946) (PMCID:PMC4780851)
    
    Coburn, Luke, Cerone, Luca, Torney, Colin ORCID: https://orcid.org/0000-0003-1673-7835, Couzin, Iain D and Neufeld, Zoltan
  
(2013)
  
Tactile interactions lead to coherent motion and enhanced chemotaxis of migrating cells.
    
    Physical Biology, 10(4),
	 
      046002.
    
    (doi: 10.1088/1478-3975/10/4/046002)
    (PMID:23752100) 
     
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Levin, Simon A. and Couzin, Iain D.
  
(2013)
  
Decision accuracy and the role of spatial interaction in opinion dynamics.
    
    Journal of Statistical Physics, 151(1-2),
	  pp. 203-217.
     
    
    (doi: 10.1007/s10955-013-0700-5)
     
     
  
  
    
    Berdahl, Andrew, Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Ioannou, Christos C., Faria, Jolyon J. and Couzin, Iain D.
  
(2013)
  
Emergent sensing of complex environments by mobile animal groups.
    
    Science, 339(6119),
	  pp. 574-576.
     
    
    (doi: 10.1126/science.1225883)
    (PMID:23372013) 
     
  
  
    
    Couzin, Iain D., Ioannou, Christos C., Demirel, Güven, Gross, Thilo, Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Hartnett, Andrew, Conradt, Larissa, Levin, Simon A. and Leonard, Naomi E.
  
(2011)
  
Uninformed individuals promote democratic consensus in animal groups.
    
    Science, 334(6062),
	  pp. 1578-1580.
     
    
    (doi: 10.1126/science.1210280)
    (PMID:22174256) 
     
  
  
    
    Torney, Colin J. ORCID: https://orcid.org/0000-0003-1673-7835, Berdahl, Andrew and Couzin, Iain D.
  
(2011)
  
Signalling and the evolution of cooperative foraging in dynamic environments.
    
    PLoS Computational Biology, 7(9),
	  e1002194.
     
    
    (doi: 10.1371/journal.pcbi.1002194)
    (PMID:21966265) 
    (PMCID:PMC3178622) 
  
  
Book Sections
    
    Campioni, Nazareno, Husmeier, Dirk ORCID: https://orcid.org/0000-0003-1673-7413, Gaskell, Jennifer 
ORCID: https://orcid.org/0000-0001-9583-323X, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490 and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2025)
  
Scalable non-Markovian state switching models for animal movement.
    
    
      In: Giuggioli, Luca and Maini, Philip (eds.)
      The Mathematics of Movement: an Interdisciplinary Approach to Mutual Challenges in Animal Ecology and Cell Biology.
    
    
    Springer Nature.
        
        
  
   (Accepted for Publication)
    
    Campioni, Nazareno, Husmeier, Dirk ORCID: https://orcid.org/0000-0003-1673-7413, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490, Gaskell, Jennifer 
ORCID: https://orcid.org/0000-0001-9583-323X and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2020)
  
Modelling multiscale collective behavior with Gaussian processes.
    
    
      In: Ladde, Gangaram and Samia, Noelle (eds.)
      Proceedings of the 2nd International Conference on Statistics: Theory and Applications (ICSTA'20).
    
    
    Avestia Publishing: Ottawa, Canada, p. 124.
         ISBN 9781927877685
        (doi: 10.11159/icsta20.124)
  
  
    
    Gaskell, Jennifer ORCID: https://orcid.org/0000-0001-9583-323X, Campioni, Nazareno, Morales, Juan M. 
ORCID: https://orcid.org/0000-0001-7269-7490, Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413 and Torney, Colin J. 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2020)
  
Approximate Bayesian inference for individual-based models with emergent dynamics.
    
    
      In: Ladde, Gangaram and Samia, Noelle (eds.)
      Proceedings of the 2nd International Conference on Statistics: Theory and Applications (ICSTA'20).
    
    
    Avestia Publishing: Ottawa, Canada, p. 125.
         ISBN 9781927877685
        (doi: 10.11159/icsta20.125)
  
  
Conference Proceedings
    
    Paun, Ionut ORCID: https://orcid.org/0009-0003-1673-9803, Husmeier, Dirk 
ORCID: https://orcid.org/0000-0003-1673-7413 and Torney, Colin 
ORCID: https://orcid.org/0000-0003-1673-7835
  
(2019)
  
A Study on Discrete-Time Movement Models.
    
    In: International Conference on Statistics: Theory and Applications (ICSTA’19), Lisbon, Portugal, 13-14 Aug 2019,
     p. 27. 
     ISBN 9781927877647
    (doi: 10.11159/icsta19.27)
  
  
Supervision
- Duffield, Christine 
 The Leverhulme Programme for Doctoral Training in Ecological Data Science
- Lindeman, Mathilde 
 The Leverhulme Programme for Doctoral Training in Ecological Data Science
- Shao, Tianle 
 The Leverhulme Programme for Doctoral Training in Ecological Data Science
- Tiddy, Izzy 
 The effects of climate-related stressors on fish social behaviour
Research datasets
2022
Kavwele, C. M., Torney, C. , Morrison, T. , Fulford, S., Masolele, M., Masoy, J. and Hopcraft, G. (2022) Non-local effects of human activity on the spatial distribution of migratory wildlife in Serengeti National Park, Tanzania. [Data Collection]
2021
Torney, C. , Morales, J. and Husmeier, D. (2021) A hierarchical machine learning framework for the analysis of large scale animal movement data. [Data Collection]
2019
Torney, C., Lloyd-Jones, D. J., Chevallier, M., Moyer, D. C., Maliti, H. T., Mwita, M., Kohi, E. M. and Hopcraft, G. (2019) A comparison of deep learning and citizen science techniques for counting wildlife in aerial survey images. [Data Collection]
2018
Torney, C., Lamont, M., Debell, L., Angohiatok, R. J., Leclerc, L.-M. and Berdahl, A. M. (2018) Inferring the rules of social interaction in migrating caribou. [Data Collection]
