School of Biodiversity, One Health & Veterinary Medicine

Dr Denise Hough

  • Lecturer (Physiology Ageing & Welfare)

telephone: 01413303705
email: Denise.Hough@glasgow.ac.uk

R235B Level 2, Jarret Building, School of Biodiversity, One Health and Veterinary Medicine, University of Glasgow, G61 1QH

Import to contacts

ORCID iDhttps://orcid.org/0000-0002-5478-693X

Biography

Dr Denise Hough is a Lecturer in Biology on the Learning, Teaching and Scholarship track at the University of Glasgow. She coordinates and teaches across undergraduate programmes in animal biology, veterinary biosciences and veterinary medicine, with a particular focus on student-centred learning and authentic assessment. Denise has led institutional and sector-wide initiatives on the responsible use of generative AI in education, including establishing the MVLS Generative AI Network for Education (GAINEd) and contributing to international Massive Online Open Courses. Alongside her pedagogical scholarship, she supervises student research projects in endocrinology and animal welfare, and is active in public engagement and widening participation.

Research interests

I am interested in how steroid hormone production changes dynamically over an individual's lifespan and the physiological processes that it influences. Some of my research are molecular and cellular based, such as investigations of tissue-specific steroid biosynthesis. Other areas of research utilise physiological and behavioural approaches to investigate the role of steroids during critical periods of development to program health later in life. 

Research groups

Publications

List by: Type | Date

Jump to: 2026 | 2025 | 2024 | 2023 | 2019 | 2018 | 2017 | 2015 | 2013 | 2012 | 2010 | 2009
Number of items: 24.

2026

Bawn, Matt, Francis, Nigel, Alvey, Elizabeth, Hassall, Christopher, Pires-daSilva, Andre, Barra, Pedro, Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Campbell, Hannah, Hardy, Matt and Canet-Perez, Juanvi (2026) Perspectives from a workshop: intelligent assessment in the age of artificial intelligence. Advances in Physiology Education, 50(1), pp. 73-82. (doi: 10.1152/advan.00246.2024)

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2026) GenAI in Assessment and Curriculum Design: AIchemy Workshop. GenAI in Chemistry Education Symposium, University of Liverpool, 21 July 2026.

2025

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2025) Top Tips and Updates from MVLS Generative AI Network for Education (GAINEd). University of Glasgow AI Symposium (UofG AI Symposium), James McCune Smith Learning Hub, University of Glasgow, Glasgow, Scotland, 10 September 2025. (doi: 10.5281/ZENODO.17304831)

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2025) RevisionBot: A Teacher-Approved AI-tool Supporting Student Self-Assessment. Anatomical Society Education Committee Webinar, Online, 5 August 2025.

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2025) Top Tips on Student Guidance for the Responsible Use of Generative AI in Higher Education. Association for Learning Technology CPD Webinar Series 2025, Online, 9 July 2025.

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Patterson, Victoria and Mable, Barbara K. ORCID logoORCID: https://orcid.org/0000-0001-7118-4116 (2025) Making research reasoning visible: redesigning a final-year literature review for a GenAI-enabled world: Published in Assessment and Feedback Case Study Compendium 2026. Advance HE Assessment and Feedback Symposium, 4 November 2025.

2024

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2024) A course coordinator's lessons learnt from the 2023-24 academic year of embracing AI in teaching practice. Intelligent Assessment in the age of AI, University of Leeds, 14 May 2024.

2023

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X and Corbishley, Alexander (2023) Guidance on responsible use of AI tools for enhanced learning – BVMS Programme. [Protocols]

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Johnstone, Iain and Gray, Joseph (2023) Life Sciences Student guidance on responsible use of AI. [Protocols]

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2023) Student guidance for the responsible use of AI. Association for Learning Technology Blog, 10 Oct.

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2023) Examples of using AI to support student learning & assessment in Life Sciences. [Website]

2019

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Robinson, J.E. ORCID logoORCID: https://orcid.org/0000-0002-4632-1366, Bellingham, M. ORCID logoORCID: https://orcid.org/0000-0002-3646-8989, Fleming, L.M., McLaughlin, M. ORCID logoORCID: https://orcid.org/0000-0001-5317-2090, Jama, K., Haraldsen, I.R.H., Solbakk, A.K. and Evans, N.P. ORCID logoORCID: https://orcid.org/0000-0001-7395-3222 (2019) Peripubertal GnRH and testosterone co-treatment leads to increased familiarity preferences in male sheep. Psychoneuroendocrinology, 108, pp. 70-77. (doi: 10.1016/j.psyneuen.2019.06.008) (PMID:31229635) (PMCID:PMC6712355)

O'Shaughnessy, P. J. et al. (2019) Alternative (backdoor) androgen production and masculinization in the human fetus. PLoS Biology, 17(2), e3000002. (doi: 10.1371/journal.pbio.3000002) (PMID:30763313) (PMCID:PMC6375548)

2018

Muvhali, Pfunzo T., Bonato, Maud, Engelbrecht, Anel, Malecki, Irek A., Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Robinson, Jane E. ORCID logoORCID: https://orcid.org/0000-0002-4632-1366, Evans, Neil P. ORCID logoORCID: https://orcid.org/0000-0001-7395-3222 and Cloete, Schalk W.P. (2018) The effect of extensive human presence at an early age on stress responses and reactivity of juvenile ostriches towards humans. Animals, 8(10), 175. (doi: 10.3390/ani8100175) (PMID:30301126) (PMCID:PMC6210211)

Johnston, Z. C. et al. (2018) The human fetal adrenal produces cortisol but no detectable aldosterone throughout the second trimester. BMC Medicine, 16, 23. (doi: 10.1186/s12916-018-1009-7) (PMID:29429410) (PMCID:PMC5808459)

2017

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Bellingham, M. ORCID logoORCID: https://orcid.org/0000-0002-3646-8989, Haraldsen, J.R., McLaughlin, M. ORCID logoORCID: https://orcid.org/0000-0001-5317-2090, Robinson, J.E. ORCID logoORCID: https://orcid.org/0000-0002-4632-1366, Solbakk, A.K. and Evans, N.P. ORCID logoORCID: https://orcid.org/0000-0001-7395-3222 (2017) A reduction in long-term spatial memory persists after discontinuation of peripubertal GnRH agonist treatment in sheep. Psychoneuroendocrinology, 77, pp. 1-8. (doi: 10.1016/j.psyneuen.2016.11.029) (PMID:27987429) (PMCID:PMC5333793)

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Bellingham, M. ORCID logoORCID: https://orcid.org/0000-0002-3646-8989, Haraldsen, I.R.H., McLaughlin, M. ORCID logoORCID: https://orcid.org/0000-0001-5317-2090, Rennie, M., Robinson, J.E. ORCID logoORCID: https://orcid.org/0000-0002-4632-1366, Solbakk, A.K. and Evans, N.P. ORCID logoORCID: https://orcid.org/0000-0001-7395-3222 (2017) Spatial memory is impaired by peripubertal GnRH agonist treatment and testosterone replacement in sheep. Psychoneuroendocrinology, 75, pp. 173-182. (doi: 10.1016/j.psyneuen.2016.10.016) (PMID:27837697) (PMCID:PMC5140006)

2015

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Storbeck, K., Cloete, S.W.P., Swart, A.C. and Swart, P. (2015) Relative contribution of P450c17 towards the acute cortisol response: lessons from sheep and goats. Molecular and Cellular Endocrinology, 408, pp. 107-113. (doi: 10.1016/j.mce.2015.01.018) (PMID:25597634)

2013

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Cloete, S.W.P., Swart, P. and Cloete, J.J.E. (2013) Searching for SNPs that affect sheep robustness: CYP17 SNP affects behavioural responses to psychological stress. In: 20th Association for the Advancement of Animal Breeding and Genetics Conference, Napier, New Zealand, 20-23 Oct 2013, pp. 503-506.

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Cloete, S.W.P., Storbeck, K., Swart, A.C. and Swart, P. (2013) Cortisol production in sheep is influenced by the functional expression of two cytochrome P450 17-alpha-hydroxylase/17,20-lyase (CYP17) isoforms. Journal of Animal Science, 91(3), pp. 1193-1206. (doi: 10.2527/jas.2012-5800)

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Swart, Pieter and Cloete, Schalk (2013) Exploration of the hypothalamic-pituitary-adrenal axis to improve animal welfare by means of genetic selection: lessons from the South African Merino. Animals, 3(2), pp. 442-474. (doi: 10.3390/ani3020442) (PMID:26487412) (PMCID:PMC4494397)

2012

Cloete, A.S., Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X and Cloete, S.W.P. (2012) Testing times – developing awareness while fostering research: improving health, welfare and production in South African Merinos. In: 2012 Australian and New Zealand Council for the Care of Animals in Research and Teaching Conference, Perth, W.A., Australia, 24 - 26 Jul 2012, pp. 43-48. ISBN 9780987465702

2010

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Cloete, S.W.P., Swart, P. and Storbeck, K. (2010) Improving stress coping ability: comparison between the CYP17 genotype Of Ovis Aries and Capra Hircus. In: 9th World Congress on Genetics Applied to Livestock Production, Leipzig, Germany, 1-6 Aug 2010, 0628.

2009

Van der Walt, D., Cloete, S.W.P., Storbeck, K. and Swart, P. (2009) The role of Cytochrome P450 17-alpha-Hydroxylase/ 17,20-Lyase (CYP17) in the stress coping ability in a divergently selected Merino sheep population. In: 18th Association for the Advancement of Animal Breeding and Genetics Conference, Barossa Valley, South Australia, 28 Sep - 1 Oct 2009, pp. 100-103.

This list was generated on Thu Sep 10 19:15:03 2026 BST.
Jump to: Articles | Conference or Workshop Item | Conference Proceedings | Protocols | Website
Number of items: 24.

Articles

Bawn, Matt, Francis, Nigel, Alvey, Elizabeth, Hassall, Christopher, Pires-daSilva, Andre, Barra, Pedro, Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Campbell, Hannah, Hardy, Matt and Canet-Perez, Juanvi (2026) Perspectives from a workshop: intelligent assessment in the age of artificial intelligence. Advances in Physiology Education, 50(1), pp. 73-82. (doi: 10.1152/advan.00246.2024)

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2023) Student guidance for the responsible use of AI. Association for Learning Technology Blog, 10 Oct.

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Robinson, J.E. ORCID logoORCID: https://orcid.org/0000-0002-4632-1366, Bellingham, M. ORCID logoORCID: https://orcid.org/0000-0002-3646-8989, Fleming, L.M., McLaughlin, M. ORCID logoORCID: https://orcid.org/0000-0001-5317-2090, Jama, K., Haraldsen, I.R.H., Solbakk, A.K. and Evans, N.P. ORCID logoORCID: https://orcid.org/0000-0001-7395-3222 (2019) Peripubertal GnRH and testosterone co-treatment leads to increased familiarity preferences in male sheep. Psychoneuroendocrinology, 108, pp. 70-77. (doi: 10.1016/j.psyneuen.2019.06.008) (PMID:31229635) (PMCID:PMC6712355)

O'Shaughnessy, P. J. et al. (2019) Alternative (backdoor) androgen production and masculinization in the human fetus. PLoS Biology, 17(2), e3000002. (doi: 10.1371/journal.pbio.3000002) (PMID:30763313) (PMCID:PMC6375548)

Muvhali, Pfunzo T., Bonato, Maud, Engelbrecht, Anel, Malecki, Irek A., Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Robinson, Jane E. ORCID logoORCID: https://orcid.org/0000-0002-4632-1366, Evans, Neil P. ORCID logoORCID: https://orcid.org/0000-0001-7395-3222 and Cloete, Schalk W.P. (2018) The effect of extensive human presence at an early age on stress responses and reactivity of juvenile ostriches towards humans. Animals, 8(10), 175. (doi: 10.3390/ani8100175) (PMID:30301126) (PMCID:PMC6210211)

Johnston, Z. C. et al. (2018) The human fetal adrenal produces cortisol but no detectable aldosterone throughout the second trimester. BMC Medicine, 16, 23. (doi: 10.1186/s12916-018-1009-7) (PMID:29429410) (PMCID:PMC5808459)

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Bellingham, M. ORCID logoORCID: https://orcid.org/0000-0002-3646-8989, Haraldsen, J.R., McLaughlin, M. ORCID logoORCID: https://orcid.org/0000-0001-5317-2090, Robinson, J.E. ORCID logoORCID: https://orcid.org/0000-0002-4632-1366, Solbakk, A.K. and Evans, N.P. ORCID logoORCID: https://orcid.org/0000-0001-7395-3222 (2017) A reduction in long-term spatial memory persists after discontinuation of peripubertal GnRH agonist treatment in sheep. Psychoneuroendocrinology, 77, pp. 1-8. (doi: 10.1016/j.psyneuen.2016.11.029) (PMID:27987429) (PMCID:PMC5333793)

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Bellingham, M. ORCID logoORCID: https://orcid.org/0000-0002-3646-8989, Haraldsen, I.R.H., McLaughlin, M. ORCID logoORCID: https://orcid.org/0000-0001-5317-2090, Rennie, M., Robinson, J.E. ORCID logoORCID: https://orcid.org/0000-0002-4632-1366, Solbakk, A.K. and Evans, N.P. ORCID logoORCID: https://orcid.org/0000-0001-7395-3222 (2017) Spatial memory is impaired by peripubertal GnRH agonist treatment and testosterone replacement in sheep. Psychoneuroendocrinology, 75, pp. 173-182. (doi: 10.1016/j.psyneuen.2016.10.016) (PMID:27837697) (PMCID:PMC5140006)

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Storbeck, K., Cloete, S.W.P., Swart, A.C. and Swart, P. (2015) Relative contribution of P450c17 towards the acute cortisol response: lessons from sheep and goats. Molecular and Cellular Endocrinology, 408, pp. 107-113. (doi: 10.1016/j.mce.2015.01.018) (PMID:25597634)

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Cloete, S.W.P., Storbeck, K., Swart, A.C. and Swart, P. (2013) Cortisol production in sheep is influenced by the functional expression of two cytochrome P450 17-alpha-hydroxylase/17,20-lyase (CYP17) isoforms. Journal of Animal Science, 91(3), pp. 1193-1206. (doi: 10.2527/jas.2012-5800)

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Swart, Pieter and Cloete, Schalk (2013) Exploration of the hypothalamic-pituitary-adrenal axis to improve animal welfare by means of genetic selection: lessons from the South African Merino. Animals, 3(2), pp. 442-474. (doi: 10.3390/ani3020442) (PMID:26487412) (PMCID:PMC4494397)

Conference or Workshop Item

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2026) GenAI in Assessment and Curriculum Design: AIchemy Workshop. GenAI in Chemistry Education Symposium, University of Liverpool, 21 July 2026.

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2025) Top Tips and Updates from MVLS Generative AI Network for Education (GAINEd). University of Glasgow AI Symposium (UofG AI Symposium), James McCune Smith Learning Hub, University of Glasgow, Glasgow, Scotland, 10 September 2025. (doi: 10.5281/ZENODO.17304831)

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2025) RevisionBot: A Teacher-Approved AI-tool Supporting Student Self-Assessment. Anatomical Society Education Committee Webinar, Online, 5 August 2025.

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2025) Top Tips on Student Guidance for the Responsible Use of Generative AI in Higher Education. Association for Learning Technology CPD Webinar Series 2025, Online, 9 July 2025.

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Patterson, Victoria and Mable, Barbara K. ORCID logoORCID: https://orcid.org/0000-0001-7118-4116 (2025) Making research reasoning visible: redesigning a final-year literature review for a GenAI-enabled world: Published in Assessment and Feedback Case Study Compendium 2026. Advance HE Assessment and Feedback Symposium, 4 November 2025.

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2024) A course coordinator's lessons learnt from the 2023-24 academic year of embracing AI in teaching practice. Intelligent Assessment in the age of AI, University of Leeds, 14 May 2024.

Conference Proceedings

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Cloete, S.W.P., Swart, P. and Cloete, J.J.E. (2013) Searching for SNPs that affect sheep robustness: CYP17 SNP affects behavioural responses to psychological stress. In: 20th Association for the Advancement of Animal Breeding and Genetics Conference, Napier, New Zealand, 20-23 Oct 2013, pp. 503-506.

Cloete, A.S., Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X and Cloete, S.W.P. (2012) Testing times – developing awareness while fostering research: improving health, welfare and production in South African Merinos. In: 2012 Australian and New Zealand Council for the Care of Animals in Research and Teaching Conference, Perth, W.A., Australia, 24 - 26 Jul 2012, pp. 43-48. ISBN 9780987465702

Hough, D. ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Cloete, S.W.P., Swart, P. and Storbeck, K. (2010) Improving stress coping ability: comparison between the CYP17 genotype Of Ovis Aries and Capra Hircus. In: 9th World Congress on Genetics Applied to Livestock Production, Leipzig, Germany, 1-6 Aug 2010, 0628.

Van der Walt, D., Cloete, S.W.P., Storbeck, K. and Swart, P. (2009) The role of Cytochrome P450 17-alpha-Hydroxylase/ 17,20-Lyase (CYP17) in the stress coping ability in a divergently selected Merino sheep population. In: 18th Association for the Advancement of Animal Breeding and Genetics Conference, Barossa Valley, South Australia, 28 Sep - 1 Oct 2009, pp. 100-103.

Protocols

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X and Corbishley, Alexander (2023) Guidance on responsible use of AI tools for enhanced learning – BVMS Programme. [Protocols]

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X, Johnstone, Iain and Gray, Joseph (2023) Life Sciences Student guidance on responsible use of AI. [Protocols]

Website

Hough, Denise ORCID logoORCID: https://orcid.org/0000-0002-5478-693X (2023) Examples of using AI to support student learning & assessment in Life Sciences. [Website]

This list was generated on Thu Sep 10 19:15:03 2026 BST.

Grants

Grants and Awards listed are those received whilst working with the University of Glasgow.

  • Is the adrenal functionally altered when puberty is delayed with GnRHa?
    Society for Endocrinology
    2017 - 2017
     
  • Investigating hypothalamic mechanisms that control LH profiles in oestrous ewes from two genetically distinct lines
    British Society for Neuroendocrinology
    2017 - 2018
     

Teaching

Digestive Physiology and Metabolism 1 (VETSCI1006) course coordinator

Animal Biology, Ecology and Evolution (BIOL2041) course coordinator

L4 Core Skills in Animal Biology (BIOL4285) course coordinator

L3 Zoology/Marine Freshwater Biology Mini-Projects and Tutorials block co-coordinator

Current Directions in Life Sciences (BIOL1018) - Antibiotic Resistance block coordinator

Fundamental Topics in Biology 2X (BIOL2039) - Deliver Genomes to Ecosystems lecturer

Veterinary Medicine (BVMS) first year Module 3: Digestion lecturer

Hons project supervision (Veterinary Biosciences and Life Sciences)

Professional activities & recognition

Prizes, awards & distinctions

  • 2026: University Team Award (University of Glasgow Teaching Excellence Awards)
  • 2025: Teaching Innovation Award Winner (LearnSci)
  • 2021: Teaching Innovation Award Finalist (LearnSci)

Professional & learned societies

  • 2013: member, British Society for Neuroendocrinology
  • 2024: MVLS representative, UofG AI in Assessment and Feedback Operational Working Group
  • 2023: Chair, UofG MVLS GAINEd Steering Group
  • 2024: SBOHVM representative, UofG MVLS TELT committee
  • 2020: panel member, UofG Academic Appeals Committee
  • 2025: Reviewer, UofG Life Sciences Misconduct Committee
  • 2026: AI Working Group, Veterinary Schools Council UK, Education Committee
  • 2022: member, Association for Learning Technology

Portrait of Dr. Denise Hough

Dr Denise Hough on Twitter